OsPLT1 negatively regulates phosphorus deficiency tolerance by modulating nitric oxide–mediated cell wall phosphorus reutilization in rice

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ID: 314285
2026
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Abstract
Phosphorus (P) deficiency severely limits rice productivity, yet the role of root developmental regulators in P homeostasis remains unclear. Here, we show that the rice PLETHORA transcription factor OsPLT1 suppresses rice tolerance to P-deficient conditions. Under P-starved conditions, osplt1 mutants accumulated more biomass and exhibited higher soluble and total P concentrations, whereas OsPLT1-overexpressing lines were hypersensitive to P deficiency. Mechanistically, osplt1 mutants displayed reduced P retention in root cell walls, particularly in the water-soluble pectin fraction, accompanied by increased water-soluble pectin uronic acid content and elevated pectin methylesterase activity. Enzymatic digestion assays confirmed enhanced release of wall-bound P in the mutants. Moreover, osplt1 mutants showed higher nitric oxide (NO) accumulation under P deficiency, whereas SNP treatment largely reduced the differences in cell wall P and soluble P between Nip and osplt1 mutants, suggesting that NO is associated with OsPLT1-dependent P reutilization. Phosphate uptake-related gene, OsPT6, was also more strongly induced in the mutants, which may contribute to improved P acquisition. Notably, OsPHR1/2 and representative PHR-responsive genes showed comparable expression among genotypes under the same P condition, suggesting that OsPLT1-mediated regulation may not primarily depend on major changes in canonical PHR signaling activity. Together, these results reveal a link between root developmental control and P homeostasis and suggest that OsPLT1 may provide a useful genetic target for enhancing phosphorus-use efficiency in rice.
Reference Key
openalex_W7161959799 Use this key to autocite in the manuscript while using SciMatic Manuscript Manager or Thesis Manager
Authors Yong Qiang Gao, Chong Wei Zhong, Yi Zhou, Li J, Jiu Huang, Liu Zheng, Ren Fang Shen, Xiao Fang Zhu
Journal Annals of botany
Year 2026
DOI
10.1093/aob/mcag142
URL
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